07/2018 — present Distinguished Professor, U.C. San Diego
07/2014 — present Presidential Chair Professor, U.C. San Diego
07/2014 — present co-director, Quantitative Biology Graduate Program, U.C. San Diego
01/2014 — 08/2015 Senior Fellow, Institute for Theoretical Studies, ETH Zurich
07/2009 — 06/2018 Professor, Section of Molecular BiologyDivision of Biological Sciences, U.C. San Diego
11/2008 — 12/2013 Distinguished Visiting Professor, Dept of Physics & Dept of Biochemistry, Hong Kong University
09/2002 — 08/2007 ChangJiang Visiting Professor, Center for Advanced Study, Tsinghua University, Beijing, China
07/1999 — 06/2018 Professor, Department of Physics,  U.C. San Diego
01/1999 — 12/1999 Visiting Professor, Center for Studies in Physics and Biology, Rockefeller University
07/1995 — 06/1999 Associate Professor, Department of Physics, U.C. San Diego
08/1994 — 06/1995 Assistant Professor, Department of Physics, State University of New York, Stony Brook
09/1993 — 08/1994 Long-term Member, School of Natural Sciences, Institute for Advanced Study, Princeton
09/1990 — 08/1993 Post-doctoral Fellow, Department of Physics, Harvard University



June 1990
Ph.D. in Physics
Massachusetts Institute of Technology, Cambridge, MA
thesis: Statistical mechanics and dynamics of surfaces and membranes
thesis advisors: Mehran Kardar and Toyoichi Tanaka
June 1986
B.S. in Physics, Biology
& Electrical Engineering
Stanford University, Stanford, CA
physics thesis : The electroweak contribution to the muon’s g-2 anomaly
EE thesis: Laser-recrystallization and silicon-on-insulator device structure


Awards & Fellowships

2020 Member, the National Academy of Sciences
2012 Fellow,  the American Academy of Microbiology
2008 Fellow, the American Physical Society
2000-03 Burroughs-Wellcome’s Innovation Award in Functional Genomics
1999-00 Guggenheim Fellowship
1997-00 Beckman Young Investigator Award
1995-98 Young Investigator AwardOffice of Naval Research
1994-99 A. P. Sloan Foundation Research Fellowship
1993 Outstanding Young Researcher Award, Overseas Chinese Physics Assoc.
1986 LeRoy Apker Award, American Physical Society

Professional Activities

2014 — present Editorial Board Member, Molecular Systems Biology
2011 — present Steering Committee Member, EcoCyc
2005 — 2011 International Advisory Committee Member, Kavli Institute for Theoretical Physics, Chinese Academy of Science
2004 — 2010 Editor-in-Chief, Physical Biology
2003 — 2009 Organizer, q-bio preprint archive
2003 — 2006 Member, Steering Committee, Kavali Institute of Theoretical Physics, Santa Barbara
2001 Organizer, Program on “Statistical Physics and Biological Information”, Kavli Institute of Theoretical Physics, Santa Barbara